The film won three other Oscars - for Best Film, Director and Best Original Screenplay.
In his acceptance speech, Firth thanked director Tom Hooper for his "immense courage".
Hooper said he had his mother to thank for finding him the King's Speech after she saw the play.
"The moral of this story is listen to your mother," he told the ceremony in Hollywood.
The film's writer, David Seidler, described how he has been moved to tears by messages thanking him for giving a voice to stutterers - of which he is one.
A heavily pregnant Natalie Portman took Best Actress for her role in Black Swan.In tears she thanked her parents and her fiance, who choreographed the film.
Welsh-born actor Christian Bale won Best Supporting Actor for The Fighter.
He said he was really touched that people thought he was worthy of his award.
Melissa Leo took the Oscar for Best Supporting Actress for her role in the same film, beating British actress Helena Bonham Carter, in The King's Speech.
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